This page contains some papers by our scientists that have appeared in peer-reviewed scientific journals as well as a selection of reports that represent important contributions to the existing knowledge.


The following publications are available:


Levine, M., Schmidt, Jennifer and others (2017)
Biology of an Endangered Marine Megavertebrate Further Informed through Citizen Science,
BioScience, bix127,
PDF Download (3.44 mb)


Levine, M., Collier, R.S., Ritter, E., Fouda, M. and Canabal, V. (2014)
Shark Cognition and a Human Mediated Driver of a Spate of Shark Attacks,
Open Journal of Animal Sciences, 4, 263-269.
PDF Download (3.44 mb)


Erich K. Ritter, Raid Amin (2013)
Are Caribbean reef sharks, Carcharhinus perezi, able to perceive human body orientation? Animal Cognition, 2013; DOI: 10.1007/s10071-013-0706-z


Erich K. Ritter (2012)
A Rare Use of a Shark’s Fin? Pectoral Fin Scooping of a Sharksucker from The Flank Area of a Female Blacktip Shark. The Open Fish Science Journal. 2012, 5, 57-59

Rain Amin, Erich Ritter (2012)
Effect of Human Body Position on the Swimming Behavior of Bull Sharks, Society & Animals, Vol. 20, No. 3, pp 225-235

Raid Amin, Erich Ritter, Peter Kennedy (2012)
A Geospatial Analysis of shark attack rates for the east coast of Florida 1994-2009. Marine and Freshwater Behaviour and Physiology: 45 (3): 185

Raid Amin, Erich Ritter, Lauren Cossette (2012)
A Geospatial Analysis of Shark Attack Rates for the Coast of California: 1994-2010, Journal of Environment and Ecology.Vol.3, No.1, pp. 246-255


Schmidt, Jennifer V, Chien-Chi Chen, Saad I. Sheikh, Mark  G. Meekan, Bradley L. Norman, Shoou-Jeng Joung (2010)
Paternity Analysis in a litter of whale shark embryos, Endangered Species Research, Vol. 12: 117-124


Schmidt, Jennifer V, Claudia L. Schmidt, Fusun Ozer, Robin E. Ernst, Kevin A. Feldheim, Mary V. Ashley, Marie Levine (2009),
Low Genetic Differentiation across Three Major Ocean Populations of the Whale Shark, Rhincodon typus. PLoS ONE 4(4): e4988. doi:10.1371/journal.pone.0004988


Levine, Marie (2008)
Whale Sharks of Donsol, Report to Explorers Club.

Potenski, Matthew, (2008) Mafia Island, Tanzania, Report of Field Research & Supplemental Work: November 2007 to March 2008.


Potenski, Matthew. (2007)
Mafia Island, Tanzania, Field Season 1: November 2006 to March 2007, Preliminary Tagging Report.

Gifford Andrew, L.J.V. Compagno, M. Levine and A. Antoniou, (2007)
Satellite tracking of whale sharks using tethered tags. Fisheries Research, Whale Sharks: Science, Conservation and Management. Proceedings of the First International Whale Shark Conference.

Gifford, Andrew, L.J.V. Compagno & Marie Levine. (2007)
Aerial Surveys of whale sharks (Rhincodon typus) off the East Coast of Southern Africa. In: T.R. Irvine and J.K. Keesing (Eds.) Proceedings of the First International Whale Shark Conference, 9-12 May 2005 Australia.

Arauz, Randall, Alex Antoniou, Ilena Zanela, L.J.V. Compagno & Marie Levine,
Preliminary Results: Census of sharks of the National Marine Park at Cocos Island: July 2005 to May 2007.


Excercise Jurassic Shark, Cocos Island July 2 – 15, 2006

Arauz, Randall & Alex Antoniou. (2006)
Preliminary Results: Movements of scalloped hammerhead sharks (Sphyrna lewini) tagged in Cocos Island National Park, Costa Rica, 2005. Memoria: Primer Seminario Taller Sobre el Estado del Conocimiento de la Condrictio fauna de Costa Rica. 8-9 (Spanish)


Ritter, Erich. K. and Marie Levine (2005),
Bite motivation of sharks reflected by the wound structure on humans. American Journal of Forensic Medicine and Pathology. Jun; 26(2):136-40.


Potenski, Matthew, (2004),
Manta Ecuador, Expedition Report

Patil, Arun. (2004) Whale Shark Project India: Feasibility Study. SRI-India.

Ritter, Erich & Marie Levine (2004)
Use of Forensic Analysis to Better Understand Shark Attack Behaviour. The Journal of Forensic Odonto-Stomatology, Vol.22 No.2, December 2004


Antoniou, Alex & Marie Levine (2003)
Archival Pop-Up Satellite Tagging of Whale Sharks (Rhincodon typus) in Honduras and the Galapagos Islands

Patil, Arun. (2002)
Whale Shark Project India: Post-Ban. SRI-India.


Patil, Arun.(2001)
Whale Shark Project India: Assessment of Whale Shark Fishery. SRI-India


Gifford, Andrew (1999)
Consolidated Summary of the fieldwork studies done on the Whale Sharks off the East Coast of Southern Africa fro the period 1993 to 1998.


Gifford, Andrew (1998)
Report on the Fifth Whale Shark Tagging and Survey Program for the period May 1, 1997 to April 30, 1998. SRI South Africa.


Gifford, Andrew (1997)
Report on the Third and Fourth Whale Shark Tagging and Survey Programs for the period May 1, 1995 to April 30, 1997. SRI South Africa.


Levine, M. (1996). Ch 40.
Unprovoked Attacks by White Sharks Off the South African Coast. The biology of Carcharodon carcharias.A. P. Klimley and D. G. Ainley. San Diego, Academic Press: 435-448


Gifford, Andrew (1994)
Preliminary whale shark tagging & survey program for the period December 1, 1993 to April 30, 1994. SRI South Africa.